In the course of investigations on the frequency of fertilization in immature rats which had been caused to ovulate by the injection of gonadotrophins, it was observed that a remarkably large number of eggs from unmated rats had divided so as to simulate segmentation very closely. Fragmentation resembling segmentation in tubal as well as ovarian eggs is a well-known phenomenon, but there is little specific information on the frequency with which it occurs.
Kampmeier [1929], who reviewed the early literature, stated that it was first reported by Pflüger in 1863 in ovarian eggs. Numerous papers have since appeared giving details of nuclear and meiotic changes observed. Kingery [1914] studied the eggs in atretic follicles of immature mice and found abnormal maturation spindles but no segmentation spindles. He frequently observed eggs with several or numerous 'cells', but these contained abnormal, several, or no nuclei. Loeb [1932] reported the finding of embryonal
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